|
|
Listado de los peces de agua dulce en la vertiente
Caribe de Venezuela (2009)
Listado y observaciones tomadas de: Rodríguez-Olarte, D., Taphorn, D. C., Lobon-Cervia,
J. 2009. Patterns of freshwater fishes of the Caribbean
versant of Venezuela. International Review of Hydrobiology, 91(1):67-90, Descargar. Premio anual de investigación UCLA 2009
El siguiente listado es producto de la
selección y valoración de registros de especies dulceacuícolas provenientes
de las colecciones de peces de CPUCLA, MCNG, MHNLS, EBRG y ICN-UNC (ver créditos al final) y de las bases de datos
de California Academy of Sciences
y FishBase. Se dispuso de referencias generales
para actualizar los registros (por ejemplo: Reis et al., 2003; Lasso et al.,
2004) y se complementó la información con inventarios locales y regionales.
Se incluyeron epítetos arbitrarios para
aquellas especies aún sin descripción taxonómica. No se incluyeron especies o
grupos periféricos que habitan en ambientes con influencia marina
(desembocaduras, estuarios) y aquellas con condición anfídroma, como las
familias Gobiidae, Ariidae y Gerreidae, entre muchas otras. En varias cuencas
de la Vertiente Caribe de Venezuela (VCV) no
existen registros completos de larga datación sobre su ictiofauna; aunado a
esto, problemas de resolución taxonómica, sugirieron excluir y/o reasignar la
distribución de algunas especies.
Registros únicos fueron considerados dudosos y se excluyeron si
conformaron hiatos geográficos o disyunciones en la distribución no
reconocidas por los especialistas.
Además, tales exclusiones se basaron también en la revisión de la
distribución reconocida y potencial para las especies, así como su condición
de tolerancia a la salinidad. Para algunas especies válidas (e.g. Hoplias teres,
Erythrinidae) no se dispusieron registros, por lo que fue asignada su
distribución sólo a la(s) cuenca(s) donde fueron reportadas inicialmente,
mientras que para otras (e.g. Caquetaia kraussii, Cichlidae) no se corroboró información que
sustente una distribución disyuntiva.
Se elaboró una matriz conteniendo
registros binarios de presencia-ausencia para 34 familias y 270 especies
dulceacuícolas de carácter primario y secundario. La condición de tolerancia
a la salinidad se asignó de acuerdo con Stiassny
& Raminosoa (1994), pero también a la
ocurrencia de las especies en estuarios y desembocaduras de los ríos al mar.
Tabla 1. Listado de los peces de agua dulce
(continentales) en la Vertiente Caribe de Venezuela y drenajes vecinos. La
tolerancia a la salinidad (T) es expresada para especies primarias (1) y
secundarias (2). Subprovincias: Magdalena: río Ranchería (M), Perijá-Andes (PA), Zulia Oriental (ZO),
Tocuyo-Yaracuy (TY), Tuy-Valencia (TV),
Neverí-Cariaco (NC), Caribe Atlántica (CA) y
Trinidad (TR).
|
Familias
|
T
|
Especies
|
MA
|
PA
|
ZO
|
TY
|
TV
|
NC
|
CA
|
TR
|
|
|
|
|
|
|
|
|
|
|
|
|
|
Ageneiosidae
|
1
|
Ageneiosus pardalis
|
|
*
|
|
|
|
|
|
|
|
Anostomidae
|
1
|
Leporinus cf friderici
|
|
|
|
|
*
|
|
|
*
|
|
Anostomidae
|
1
|
Leporinus muyscorum
|
*
|
|
|
|
|
|
|
|
|
Anostomidae
|
1
|
Leporinus cf striatus
|
*
|
|
|
|
|
|
|
|
|
Anostomidae
|
1
|
Schizodon corti
|
|
*
|
|
|
|
|
|
|
|
Apteronotidae
|
1
|
Apteronotus leptorhynchus
|
|
*
|
|
|
|
|
|
|
|
Apteronotidae
|
1
|
Apteronotus electrosensilis
|
|
|
|
*
|
|
|
|
|
|
Apteronotidae
|
1
|
Apteronotus cuchillo
|
|
*
|
*
|
|
|
|
|
|
|
Apteronotidae
|
1
|
Apteronotus cuchillejo
|
|
*
|
|
|
|
|
|
|
|
Aspredinidae
|
1
|
Aspredinichthys filamentosus
|
|
|
|
|
|
|
*
|
*
|
|
Aspredinidae
|
1
|
Aspredinichthys tibicen
|
|
|
|
|
|
|
*
|
*
|
|
Aspredinidae
|
1
|
Aspredo aspredo
|
|
|
|
|
|
|
*
|
*
|
|
Aspredinidae
|
1
|
Bunocephalus amaurus
|
|
|
|
|
*
|
*
|
|
|
|
Aspredinidae
|
1
|
Dupouyichthys sapito
|
|
*
|
|
|
|
|
|
|
|
Aspredinidae
|
1
|
Hoplomyzon atrizona
|
|
*
|
|
|
|
|
|
|
|
Aspredinidae
|
1
|
Platystacus cotylephorus
|
|
|
|
|
|
|
*
|
*
|
|
Aspredinidae
|
1
|
Xiliphius kryptos
|
|
*
|
|
|
|
|
|
|
|
Astroblepidae
|
1
|
Astroblepus chotae
|
|
*
|
|
|
|
|
|
|
|
Astroblepidae
|
1
|
Astroblepus orientalis
|
|
*
|
|
|
|
|
|
|
|
Astroblepidae
|
1
|
Astroblepus phelpsi
|
|
*
|
|
|
|
|
|
|
|
Atherinopsidae
|
1
|
Atherinella venezuelae
|
|
|
|
|
*
|
|
|
|
|
Auchenipteridae
|
1
|
Trachelyopterus peloichthys
|
|
*
|
*
|
|
|
|
|
|
|
Auchenipteridae
|
2
|
Pseudauchenipterus nodosus
|
|
|
|
|
|
|
*
|
*
|
|
Auchenipteridae
|
2
|
Trachelyopterus galeatus
|
|
|
|
|
*
|
|
|
*
|
|
Auchenipteridae
|
1
|
Trachelyopterus insignis
|
*
|
|
|
|
|
|
|
|
|
Callichthyidae
|
2
|
Callichthys callichthys
|
|
|
|
|
|
|
*
|
*
|
|
Callichthyidae
|
1
|
Corydoras aeneus
|
|
|
|
*
|
*
|
|
|
*
|
|
Callichthyidae
|
2
|
Hoplosternum littorale
|
|
|
|
|
*
|
|
*
|
*
|
|
Callichthyidae
|
2
|
Hoplosternum magdalenae
|
|
*
|
*
|
*
|
|
|
|
|
|
Callichthyidae
|
2
|
Megalechis thoracata
|
|
*
|
*
|
*
|
|
|
|
*
|
|
Cetopsidae
|
1
|
Pseudocetopsis motatanensis
|
|
*
|
|
|
|
|
|
|
|
Cetopsidae
|
1
|
Pseudocetopsis orinoco
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Aphyocharax alburnus
|
|
|
|
|
*
|
*
|
|
|
|
Characidae
|
1
|
Astyanax bimaculatus
|
|
|
|
*
|
*
|
*
|
*
|
*
|
|
Characidae
|
1
|
Astyanax cf caucanus
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
2
|
Astyanax fasciatus
|
*
|
*
|
*
|
*
|
*
|
|
|
|
|
Characidae
|
2
|
Astyanax magdalenae
|
|
*
|
*
|
|
|
|
|
|
|
Characidae
|
1
|
Astyanax metae
|
|
|
|
*
|
*
|
*
|
*
|
|
|
Characidae
|
1
|
Astyanax venezuelae
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Brycon moorei
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
1
|
Bryconamericus alpha
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Bryconamericus loisae
|
|
*
|
*
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Bryconamericus beta
|
|
|
|
|
|
|
*
|
|
|
Characidae
|
1
|
Bryconamericus cismontanus
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Bryconamericus charalae
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Bryconamericus deuterodonoides
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Knodus meridae
|
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Bryconamericus motatanensis
|
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Bryconamericus sp Alto Tocuyo
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Bryconamericus yokiae
|
|
|
|
|
*
|
|
|
|
|
Characidae
|
1
|
Bryconamericus lassorum
|
|
|
|
|
|
|
*
|
|
|
Characidae
|
1
|
Cheirodon pulcher
|
|
|
|
*
|
*
|
|
*
|
*
|
|
Characidae
|
1
|
Cheirodontops geayi
|
|
|
|
|
*
|
|
|
|
|
Characidae
|
1
|
Corynopoma riisei
|
|
*
|
|
|
*
|
*
|
|
*
|
|
Characidae
|
1
|
Creagrutus affinis
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus bolivari
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Creagrutus crenatus
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus hildebrandi
|
|
*
|
*
|
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus hysginus
|
|
|
|
|
|
*
|
*
|
|
|
Characidae
|
1
|
Creagrutus lassoi
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus lepidus
|
|
|
|
*
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus maracaiboensis
|
*
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus melasma
|
|
|
|
*
|
*
|
*
|
*
|
|
|
Characidae
|
1
|
Creagrutus paralacus
|
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Creagrutus taphorni
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Ctenobrycon spilurus
|
|
|
|
|
*
|
|
*
|
|
|
Characidae
|
1
|
Cynopotamus venezuelae
|
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Gephyrocharax melanocheir
|
*
|
|
*
|
*
|
|
|
|
|
|
Characidae
|
1
|
Gephyrocharax valencia
|
|
|
|
*
|
*
|
*
|
*
|
|
|
Characidae
|
1
|
Gephyrocharax venezuelae
|
|
*
|
*
|
*
|
|
|
|
|
|
Characidae
|
1
|
Gilbertolus maracaiboensis
|
|
*
|
*
|
|
|
|
|
|
|
Characidae
|
1
|
Gymnocorymbus thayeri
|
|
|
|
|
|
*
|
*
|
*
|
|
Characidae
|
1
|
Hemibrycon guppyi
|
|
|
|
|
|
|
|
*
|
|
Characidae
|
1
|
Hemibrycon jabonero
|
*
|
*
|
*
|
*
|
|
|
|
|
|
Characidae
|
1
|
Hemibrycon metae
|
|
|
|
*
|
*
|
*
|
*
|
|
|
Characidae
|
1
|
Hemibrycon taeniurus
|
|
|
|
|
|
*
|
|
*
|
|
Characidae
|
1
|
Hemigrammus unilineatus
|
|
|
|
|
|
|
*
|
*
|
|
Characidae
|
1
|
Hemigrammus unitaeniatus
|
|
|
|
|
|
|
|
*
|
|
Characidae
|
1
|
Hyphessobrycon cf axelrodi
|
|
|
|
|
|
|
*
|
*
|
|
Characidae
|
1
|
Hyphessobrycon bentosi
|
|
|
|
|
|
|
*
|
|
|
Characidae
|
2
|
Hyphessobrycon fernandezi
|
|
|
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Hyphessobrycon cf inconstans
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
2
|
Hyphessobrycon sovichthys
|
|
*
|
*
|
*
|
*
|
|
|
|
|
Characidae
|
1
|
Markiana geayi
|
|
|
|
|
|
|
*
|
|
|
Characidae
|
1
|
Moenkhausia pittieri
|
|
|
|
|
*
|
|
|
|
|
Characidae
|
1
|
Mylossoma acanthogaster
|
|
*
|
|
|
|
|
|
|
|
Characidae
|
1
|
Nanocheirodon insignis
|
|
*
|
*
|
*
|
|
|
|
|
|
Characidae
|
1
|
Paragoniates alburnus
|
|
|
|
|
|
|
*
|
|
|
Characidae
|
1
|
Phenagoniates macrolepis
|
|
*
|
*
|
|
|
|
|
|
|
Characidae
|
2
|
Roeboides dayi
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
2
|
Roeboides dientonito
|
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
|
Characidae
|
1
|
Saccoderma melanostigma
|
*
|
*
|
*
|
|
|
|
|
|
|
Characidae
|
1
|
Salminus affinis
|
*
|
|
|
|
|
|
|
|
|
Characidae
|
1
|
Serrasalmus neveriensis
|
|
|
|
|
*
|
*
|
|
|
|
Characidae
|
1
|
Triportheus elongatus
|
|
|
|
|
|
|
*
|
*
|
|
Characidae
|
1
|
Xenagoniates bondi
|
|
|
|
|
|
|
*
|
|
|
Cichlidae
|
2
|
Aequidens latifrons
|
*
|
|
|
|
|
|
|
|
|
Cichlidae
|
2
|
Andinoacara pulcher
|
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
|
Cichlidae
|
2
|
Apistogramma guttata
|
|
|
|
|
|
|
*
|
|
|
Cichlidae
|
2
|
Apistogramma hoignei
|
|
|
|
|
|
|
*
|
|
|
Cichlidae
|
2
|
Astronotus sp
|
|
|
|
|
|
|
*
|
|
|
Cichlidae
|
2
|
Caquetaia kraussii
|
|
*
|
*
|
|
|
|
|
|
|
Cichlidae
|
2
|
Caquetaia umbrifera
|
*
|
|
|
|
|
|
|
|
|
Cichlidae
|
2
|
Cichlasoma orinocense
|
|
|
|
|
*
|
|
*
|
|
|
Cichlidae
|
2
|
Cichlasoma taenia
|
|
|
|
|
*
|
*
|
*
|
*
|
|
Cichlidae
|
2
|
Cleithracara maronii
|
|
|
|
|
|
|
*
|
*
|
|
Cichlidae
|
2
|
Crenicichla frenata
|
|
|
|
|
|
*
|
*
|
*
|
|
Cichlidae
|
2
|
Crenicichla geayi
|
|
|
|
*
|
*
|
*
|
*
|
|
|
Cichlidae
|
2
|
Crenicichla saxatilis
|
|
|
|
|
|
*
|
*
|
*
|
|
Cichlidae
|
2
|
Geophagus steindachneri
|
*
|
*
|
|
|
|
|
|
|
|
Crenuchidae
|
1
|
Characidium boaevistae
|
|
*
|
|
|
|
|
|
|
|
Crenuchidae
|
1
|
Characidium chupa
|
*
|
*
|
|
*
|
*
|
*
|
*
|
|
|
Ctenoluciidae
|
2
|
Ctenolucius hujeta
|
*
|
*
|
*
|
|
|
|
|
|
|
Curimatidae
|
1
|
Curimata cyprinoides
|
|
|
|
|
|
|
*
|
|
|
Curimatidae
|
2
|
Cyphocharax aspilos
|
|
*
|
*
|
|
|
|
|
|
|
Curimatidae
|
1
|
Cyphocharax magdalenae
|
*
|
|
|
|
|
|
|
|
|
Curimatidae
|
1
|
Cyphocharax spilurus
|
|
|
|
|
*
|
|
*
|
|
|
Curimatidae
|
2
|
Potamorhina laticeps
|
|
*
|
|
|
|
|
|
|
|
Curimatidae
|
1
|
Steindachnerina argentea
|
|
|
|
*
|
*
|
*
|
|
*
|
|
Doradidae
|
1
|
Doraops zuloagai
|
|
*
|
|
|
|
|
|
|
|
Doradidae
|
1
|
Rhinodoras thomersoni
|
|
*
|
|
|
|
|
|
|
|
Erythrinidae
|
1
|
Erythrinus erythrinus
|
|
|
|
|
|
|
*
|
*
|
|
Erythrinidae
|
1
|
Hoplerythrinus cinereus
|
|
|
|
|
|
|
|
*
|
|
Erythrinidae
|
1
|
Hoplerythrinus unitaeniatus
|
|
|
|
|
|
|
*
|
*
|
|
Erythrinidae
|
2
|
Hoplias malabaricus
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
|
Erythrinidae
|
1
|
Hoplias teres
|
|
*
|
|
|
|
|
|
|
|
Gasteropelecidae
|
1
|
Gasteropelecus maculatus
|
*
|
*
|
*
|
|
|
|
|
|
|
Gasteropelecidae
|
1
|
Gasteropelecus sternicla
|
|
|
|
|
|
|
*
|
*
|
|
Gymnotidae
|
1
|
Gymnotus carapo
|
|
|
|
|
*
|
|
*
|
*
|
|
Heptapteridae
|
1
|
Cetopsorhamdia picklei
|
|
*
|
|
|
|
|
|
|
|
Heptapteridae
|
1
|
Cetopsorhamdia sp
|
|
|
|
*
|
|
|
|
|
|
Heptapteridae
|
1
|
Cetopsorhamdia sp Rancheria
|
*
|
|
|
|
|
|
|
|
|
Heptapteridae
|
1
|
Imparfinis nemacheir
|
|
*
|
|
|
|
|
|
|
|
Heptapteridae
|
1
|
Pimelodella cristata
|
|
|
|
*
|
*
|
|
*
|
|
|
Heptapteridae
|
2
|
Pimelodella odynea
|
|
*
|
*
|
*
|
|
|
|
|
|
Heptapteridae
|
1
|
Pimelodella sp Rancheria
|
*
|
|
|
|
|
|
|
|
|
Heptapteridae
|
1
|
Pimelodella tapatapae
|
|
|
|
|
*
|
|
|
|
|
Heptapteridae
|
1
|
Rhamdella sp San Juan
|
|
|
|
|
|
|
*
|
|
|
Heptapteridae
|
1
|
Rhamdia foina
|
|
|
|
|
|
|
*
|
|
|
Heptapteridae
|
1
|
Rhamdia humilis
|
|
|
|
|
*
|
|
|
|
|
Heptapteridae
|
1
|
Rhamdia quelen
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
|
Hypopomidae
|
1
|
Brachyhypopomus diazi
|
|
|
|
*
|
|
|
|
|
|
Hypopomidae
|
1
|
Brachyhypopomus occidentalis
|
|
*
|
*
|
|
|
|
|
|
|
Lebiasinidae
|
1
|
Piabucina erythrinoides
|
|
*
|
*
|
*
|
|
|
|
|
|
Lebiasinidae
|
1
|
Pyrrhulina lugubris
|
|
|
|
|
|
|
*
|
|
|
Loricariidae
|
1
|
Ancistrus brevifilis
|
|
*
|
*
|
*
|
*
|
*
|
*
|
|
|
Loricariidae
|
1
|
Ancistrus gymnorhynchus
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Ancistrus nationi
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Ancistrus triradiatus
|
|
*
|
|
*
|
|
|
*
|
|
|
Loricariidae
|
1
|
Ancistrus sp
Trinidad
|
|
|
|
|
|
|
|
*
|
|
Loricariidae
|
1
|
Chaetostoma anomalum
|
|
*
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma guairense
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma machiquense
|
|
*
|
*
|
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma milesi
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma nudirostre
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma pearsei
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma sovichthys
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma stanni
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma tachiraense
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma venezuelae
|
|
|
|
|
|
|
*
|
|
|
Loricariidae
|
1
|
Chaetostoma yurubiense
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Chaetostoma sp
Trinidad
|
|
|
|
|
|
|
|
*
|
|
Loricariidae
|
1
|
Cordylancistrus nephelion
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Cordylancistrus perijae
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Cordylancistrus sp Rancheria
|
*
|
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Crossoloricaria venezuelae
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Dasyloricaria filamentosa
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
2
|
Dolichancistrus cobrensis
|
|
*
|
*
|
|
|
|
|
|
|
Loricariidae
|
1
|
Farlowella acus
|
|
|
|
*
|
*
|
|
|
|
|
Loricariidae
|
1
|
Farlowella curtirostra
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Farlowella mariaelenae
|
|
|
|
*
|
*
|
|
|
|
|
Loricariidae
|
1
|
Farlowella martini
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Farlowella taphorni
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Farlowella venezuelensis
|
|
|
|
|
|
|
*
|
|
|
Loricariidae
|
1
|
Farlowella vittata
|
|
|
|
|
*
|
*
|
*
|
|
|
Loricariidae
|
1
|
Hemiancistrus maracaiboensis
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Hypostomus emarginatus
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Hypostomus hondae
|
*
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Hypostomus pagei
|
|
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Hypostomus plecostomoides
|
|
|
|
|
*
|
|
*
|
|
|
Loricariidae
|
1
|
Hypostomus plecostomus
|
|
|
|
*
|
*
|
|
*
|
|
|
Loricariidae
|
1
|
Hypostomus robinii
|
|
|
|
|
|
|
|
*
|
|
Loricariidae
|
1
|
Hypostomus villarsi
|
|
|
|
|
|
|
|
|
|
Loricariidae
|
2
|
Hypostomus watwata
|
|
*
|
*
|
*
|
|
*
|
*
|
|
|
Loricariidae
|
1
|
Lamontichthys maracaibero
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Lasiancistrus guacharote
|
|
*
|
*
|
|
|
|
|
|
|
Loricariidae
|
1
|
Lithogenes valencia
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Loricaria cataphracta
|
|
|
|
|
|
|
*
|
|
|
Loricariidae
|
1
|
Loricariichthys brunneus
|
|
|
|
|
*
|
*
|
|
|
|
Loricariidae
|
1
|
Pseudancistrus sp Tuy
|
|
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Pterygoplichthys multiradiatus
|
|
|
|
|
|
|
*
|
|
|
Loricariidae
|
1
|
Pterygoplichthys zuliaensis
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Rineloricaria caracasensis
|
|
*
|
|
|
*
|
|
|
|
|
Loricariidae
|
1
|
Rineloricaria magdalenae
|
*
|
*
|
*
|
|
|
|
|
|
|
Loricariidae
|
1
|
Rineloricaria rupestris
|
|
*
|
|
*
|
|
|
|
|
|
Loricariidae
|
1
|
Spatuloricaria lagoichthys
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Spatuloricaria phelpsi
|
|
*
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Sturisoma aureum
|
*
|
|
|
|
|
|
|
|
|
Loricariidae
|
1
|
Sturisoma kneri
|
|
*
|
*
|
|
|
|
|
|
|
Loricariidae
|
1
|
Sturisomatichthys leightoni
|
|
*
|
|
|
|
|
|
|
|
Nandidae
|
1
|
Polycentrus schomburgki
|
|
|
|
|
|
|
*
|
*
|
|
Parodontidae
|
1
|
Parodon apolinari
|
|
|
|
*
|
|
|
|
|
|
Parodontidae
|
1
|
Parodon suborbitale
|
*
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Brachyplatystoma vaillantii
|
|
|
|
|
|
|
*
|
*
|
|
Pimelodidae
|
2
|
Cheirocerus abuelo
|
|
*
|
*
|
|
|
|
|
|
|
Pimelodidae
|
2
|
Hypophthalmus marginatus
|
|
|
|
|
|
|
*
|
|
|
Pimelodidae
|
1
|
Megalonema psammium
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Perrunichthys perruno
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Pimelodus blochii
|
|
|
|
|
|
|
*
|
|
|
Pimelodidae
|
2
|
Pimelodus coprophagus
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Pimelodus grosskopfii
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Pimelodus navarroi
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Platysilurus malarmo
|
|
*
|
|
|
|
|
|
|
|
Pimelodidae
|
1
|
Sorubim cuspicaudus
|
|
*
|
*
|
|
|
|
|
|
|
Poeciliidae
|
2
|
Micropoecilia picta
|
|
|
|
|
*
|
|
*
|
*
|
|
Poeciliidae
|
2
|
Poecilia boesemani
|
|
|
|
|
|
|
|
*
|
|
Poeciliidae
|
2
|
Poecilia caucana
|
*
|
*
|
*
|
*
|
*
|
|
|
|
|
Poeciliidae
|
2
|
Poecilia dauli
|
|
|
|
*
|
|
|
|
|
|
Poeciliidae
|
2
|
Poecilia koperi
|
|
|
|
*
|
*
|
|
|
|
|
Poeciliidae
|
2
|
Poecilia parae
|
|
|
|
|
|
|
*
|
|
|
Poeciliidae
|
2
|
Poecilia reticulata
|
|
*
|
|
*
|
*
|
*
|
*
|
*
|
|
Poeciliidae
|
2
|
Poecilia sphenops
|
|
|
|
*
|
*
|
*
|
|
|
|
Poeciliidae
|
2
|
Poecilia vivipara
|
|
|
|
*
|
*
|
|
*
|
*
|
|
Poeciliidae
|
2
|
Pseudolimia heterandria
|
|
|
*
|
*
|
*
|
|
|
|
|
Poeciliidae
|
2
|
Tomeurus gracilis
|
|
|
|
|
*
|
|
*
|
|
|
Potamotrygonidae
|
1
|
Potamotrygon yepezi
|
|
*
|
|
|
|
|
|
|
|
Potamotrygonidae
|
2
|
Potamotrygon orbignyi
|
|
|
|
|
|
|
*
|
|
|
Prochilodontidae
|
1
|
Ichthyoelephas longirostris
|
*
|
|
|
|
|
|
|
|
|
Prochilodontidae
|
2
|
Prochilodus mariae
|
|
|
|
|
|
|
*
|
|
|
Prochilodontidae
|
1
|
Prochilodus reticulatus
|
*
|
*
|
*
|
|
|
|
|
|
|
Pseudopimelodidae
|
1
|
Batrochoglanis acanthochiroides
|
|
*
|
|
|
|
|
|
|
|
Pseudopimelodidae
|
1
|
Microglanis iheringi
|
|
|
|
|
*
|
|
*
|
|
|
Pseudopimelodidae
|
1
|
Microglanis poecilus
|
|
|
|
|
|
|
*
|
|
|
Pseudopimelodidae
|
1
|
Pseudopimelodus sp Maracaibo
|
*
|
*
|
|
|
|
|
|
|
|
Pseudopimelodidae
|
1
|
Pseudopimelodus mathisoni
|
|
|
|
*
|
|
|
|
|
|
Pseudopimelodidae
|
1
|
Pseudopimelodus bufonius
|
|
|
|
|
|
|
*
|
|
|
Rivulidae
|
2
|
Austrofundulus goajira
|
|
|
|
|
|
|
|
|
|
Rivulidae
|
2
|
Austrofundulus leohoignei
|
|
|
|
*
|
|
|
|
|
|
Rivulidae
|
2
|
Austrofundulus leoni
|
|
*
|
*
|
|
|
|
|
|
|
Rivulidae
|
2
|
Austrofundulus limnaeus
|
|
*
|
*
|
|
|
|
|
|
|
Rivulidae
|
2
|
Austrofundulus transilis
|
|
|
|
|
*
|
|
|
|
|
Rivulidae
|
2
|
Gnatholebias zonatus
|
|
|
|
|
*
|
|
*
|
|
|
Rivulidae
|
2
|
Rachovia brevis
|
|
|
|
|
|
|
|
|
|
Rivulidae
|
2
|
Rachovia hummelincki
|
|
|
*
|
|
|
|
|
|
|
Rivulidae
|
2
|
Rachovia maculipinnis
|
|
|
|
|
*
|
|
|
|
|
Rivulidae
|
2
|
Rachovia pyropunctata
|
|
*
|
*
|
|
|
|
|
|
|
Rivulidae
|
2
|
Rivulus deltaphilus
|
|
|
|
|
|
|
*
|
|
|
Rivulidae
|
2
|
Rivulus hartii
|
|
|
|
*
|
*
|
*
|
*
|
*
|
|
Rivulidae
|
2
|
Rivulus marmoratus
|
|
|
|
|
|
|
|
|
|
Sciaenidae
|
1
|
Pachypops fourcroi
|
|
|
|
|
|
|
*
|
|
|
Sciaenidae
|
1
|
Plagioscion magdalenae
|
*
|
|
|
|
|
|
|
|
|
Sciaenidae
|
1
|
Plagioscion squamosissimus
|
|
|
|
|
|
|
*
|
|
|
Sciaenidae
|
1
|
Plagioscion auratus
|
|
|
|
|
|
|
*
|
|
|
Sternopygidae
|
1
|
Distocyclus goajira
|
|
*
|
|
|
|
|
|
|
|
Sternopygidae
|
1
|
Eigenmannia virescens
|
*
|
*
|
*
|
|
*
|
|
|
|
|
Sternopygidae
|
1
|
Sternopygus macrurus
|
|
*
|
|
|
*
|
|
*
|
|
|
Sternopygidae
|
1
|
Sternopygus pejeraton
|
*
|
*
|
*
|
|
|
|
|
|
|
Synbranchidae
|
2
|
Ophisternon aenigmaticum
|
|
|
|
|
|
|
|
*
|
|
Synbranchidae
|
2
|
Synbranchus marmoratus
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
*
|
|
Trichomycteridae
|
1
|
Ochmacanthus cf alternus
|
|
|
|
|
*
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus arleoi
|
|
|
|
*
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus emanuelli
|
|
*
|
|
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus cf bogotense
|
|
|
|
*
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus maracaiboensis
|
|
*
|
|
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus meridae
|
|
*
|
|
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus mondolfi
|
|
|
|
|
*
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus spelaeus
|
|
|
|
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus sp Tocuyo
|
|
|
|
*
|
|
|
|
|
|
Trichomycteridae
|
1
|
Trichomycterus sp San Juan
|
|
|
|
|
|
|
*
|
|
|
Trichomycteridae
|
1
|
Tridensimilis venezuelae
|
|
*
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
Agradecemos a las siguientes Colecciones
biológicas y su personal asociado por el acceso a sus bases de datos: CPUCLA,
Colección Regional de Peces UCLA (Douglas Rodríguez Olarte). MCNG, Museo de
Ciencias Naturales Guanare UNELLEZ (Donald C. Taphorn, Otto Castillo, Críspulo
Marrero). MHNLS, Museo de Historia
Natural La Salle FLASA (Carlos Lasso, Oscar Lasso) EBRG, Estación Biológica de Rancho Grande MPPAMB (Marcos Campo). MBUCV,
Museo de Biología UCV (Francisco Provenzano). ICN, Instituto de Ciencias Naturales UNC
(José Iván Mojica).
|
|
